Abdel-Rahman, M.S., Fahmy, M.M. and Aggour, M., 1998. Trials for control of ixodid ticks using pheromone acaricide tick decoys. Journal of Egyptian Society of Parasitology 28: 551-557. Google Scholar
	Allan, S.A., 2010. Chemical ecology of tick-host interactions. In: Takken, W. and Knols, B.G.J. (eds.) Olfaction in vector-host interactions, Wageningen Academic Publishers, Wageningen, the Netherlands, pp. 327-348. Google Scholar
	Allan, S.A. and Sonenshine, D.E., 2002. Evidence of an assembly pheromone in the black-legged deer tick, Ixodes scapularis. Journal of Chemical Ecology 28: 15-27. https://doi.org/10.1023/A:1013554517148 Crossref, Google Scholar
	Allan, S.A., Barré, N., Sonenshine, D.E., Burridge, M.J., 1998. Efficacy of tags impregnated with pheromone and acaricide for control of Amblyomma variegatum. Medical and Veterinary Entomology 12: 141-150. https://doi.org/10.1046/j.1365-2915.1998.00080.x Crossref, Google Scholar
	Allan, S.A., Phillips, J.S. and Sonenshine, D.E., 1989. Species recognition elicited by differences in composition of the genital sex pheromone in Dermacentor variabilis and D. andersoni (Acari: Ixodidae). Journal of Medical Entomology 26: 539-546. https://doi.org/10.1093/jmedent/26.6.539 Crossref, Google Scholar
	Allan, S.A., Phillips, J.S. and Sonenshine, D.E., 1991. Role of genital sex pheromones in Amblyomma americanum and A. maculatum (Acari: Ixodidae). Experimental and Applied Acarology 11: 9-21. https://doi.org/10.1007/BF01193725 Crossref, Google Scholar
	Allan, S.A., Phillips, J.S., Taylor, D. and Sonenshine, D.E., 1988. Genital sex pheromones of ixodid ticks: evidence for the role of fatty acids from the anterior reproductive tract in mating of Dermacentor variabilis and Dermacentor andersoni. Journal of Insect Physiology 34: 315-323. https://doi.org/10.1016/0022-1910(88)90142-4 Crossref, Google Scholar
	Allan, S.A., Sonenshine, S.A. and Burridge, M.J., 2001. Tick pheromones and uses thereof. U.S. Patent No. 6331297. Andrews, R.H. and Bull, C.M., 1980. Mating behaviour in the Australian reptile tick Aponomma hydrosauri. Animal Behaviour 28: 1280-IN1. https://doi.org/10.1016/S0003-3472(80)80116-3 Crossref, Google Scholar
	Anish, R.K., Latha, B.R., Ramanathan, G., Sivagnanam, U.T., Sreekumar, C. and Leela, V., 2017. A novel assembly pheromone trap for tick control in dog kennels. Veterinary Parasitology 235: 57-63. https://doi.org/10.1016/j.vetpar.2017.01.005 Crossref, Google Scholar
	Apps, P.J., Viljoen, H.W. and Pretorius, V., 1988. Aggregation pheromones of the bont tick Amblyomma hebraeum: identification of candidates for bioassay. Onderstepoort Journal of Veterinary Research 55: 135-137. http://hdl.handle.net/2263/42374 Google Scholar
	Barré, N., Aprelon, R. and Eugène, M., 1998. Attempts to feed Amblyomma variegatum ticks on artificial membranes. Annals of the New York Academy of Science 849: 384-390. https://doi.org/10.1111/j.1749-6632.1998.tb11077.x Crossref, Google Scholar
	Barré, N., Garris, G.I. and Lorvelec, O., 1997. Field sampling of the tick Amblyomma variegatum (Acari: Ixodidae) on pastures in Guadeloupe; attraction of CO2 and/or tick pheromones and conditions of use. Experimental and Applied Acarology 21: 95-108. https://doi.org/10.1023/B:APPA.0000031788.88306.77 Crossref, Google Scholar
	Berger, R.S., 1972. 2,6-dichlorophenol, sex pheromone of the lone star tick. Science 177: 704-705. Crossref, Google Scholar
	Benoit, J.B., Yoder, J.A., Lopez-Martinez, G., Elnitsky, M.A., Lee, R.E. Jr. and Denlinger, D.L. 2007. Habitat requirement of the seabird tick, Ixodes uriae (Acari: Ixodidae), from the Antarctic Peninsula in relation to water balance characteristics of eggs, non-fed and engorged stages. Journal of Comparative Physiology B 177: 205-215. https://doi10.1007/s00360-006-0122-7 Crossref, Google Scholar
	Benoit, J.B., Lopez-Martinez, G., Philips, S.A., Elnitsky, M.A., Yoder, J.A., Lee, R.E. and Denlinger, D.L., 2008. The seabird tick, Ixodes uriae, uses uric acid in penguin guano as a kairomone and guanine in tick feces as an assembly pheromone on the Antarctic Peninsula. Polar Biology 31: 1445-1451. https://doi.org/10.1007/s00300-008-0485-1 Crossref, Google Scholar
	Bhoopathy, D. and Latha, B.R., 2018. An eco-friendly approach to control Rhipicephalus sanguineus by using sustained release porous calcium alginate beans containing assembly pheromone. Exploratory Animal and Medical Research 8: 33-39. Google Scholar
	Bhoopathy, D. and Ravi, L.B., 2017. In vitro trials to ascertain sustained release efficacy of assembly pheromone microparticles for the control of brown dog tick, Rhipicephalus sanguineus. Journal of Parasitic Diseases 41: 11431146. Crossref, Google Scholar
	Bhoopathy, D., Latha, B., Sreekumar, C. and Leela, V., 2016. Response of unfed stages Rhipicephalus sanguineus to subtle variations in the composition of assembly pheromone. Journal of Parasitic Diseases 40: 1392-1395. Crossref, Google Scholar
	Bhoopathy, D., Latha, B.R., Uma, T.S., Sreekumar, C. and Leela, V., 2014. A novel approach to control brown dog tick, Rhipicephalus sanguineus using sustained release poly^-caprolactone-pheromone microspheres. Acta Parasitología 59: 153-157. Crossref, Google Scholar
	Borges, L.M.F. and Ribeiro, M.F.B., 1999. Presence of sex pheromones in Anocentor nitens (Acari: Ixodidae). Journal of Medical Entomology 36: 50-54. https://doi.org/10.1093/jmedent/36.L50 Google Scholar
	Borges, L.M., Eiras, Á.E., Ferri, P.H. and Lôbo, A.C., 2002. The role of 2, 6-dichlorophenol as sex pheromone of the tropical horse tick Anocentor nitens (Acari: Ixodidae). Experimental and Applied Acarology 27: 223-230. https://doi.org/10.1023/A:1021620215416 Crossref, Google Scholar
	Borges, L.M.F., Ferreira, L.A.M., da Silva, L.S., de Oliveira, R.A., Mussi, S.V., Faria, K.A., Melo, L.S., Abud, L.J., Costa, G.L and Soares, S.F., 2007. Efficacy of 2, 6-dichlorophenol lure to control Dermacentor nitens. Veterinary Parasitology 147: 155-160. https://doi.org/10.1016/j.vetpar.2007.02.037 Crossref, Google Scholar
	Bouman, E.A., Dusbábek, F., Simek, P. and Zahradniekova, H., 2003. Methyl 3-chloro-4-methoxybenzoate, a new candidate semiochemical inhibiting copulation behaviour of Ixodes ricinus (L.) males. Physiological Entomology 28: 276-282. https://doi.org/10.1111/j.1365-3032.2003.00343.x Crossref, Google Scholar
	Bryson, N.R., Horak, I.G., Venter, E.H. and Yunker, C.E., 2000. Collection of free-living nymphs and adults of Amblyomma hebraeum (Acari: Ixodidae) with pheromone/carbon dioxide traps at 5 different ecological sites in heartwater endemic regions of South Africa. Experimental and Applied Acarology 24: 971-982. https://doi.org/10.1023/A:1010639113793 Crossref, Google Scholar
	Carnohan, L.P., Kaufman, P.E., Allan, S.A., Gezan, S.A. and Weeks, E.N., 2017. Laboratory and field evaluation of brown dog tick behavioural responses to potential semiochemicals. Ticks and Tick-borne Diseases 8: 226-234. https://doi.org/10.1016/j.ttbdis.2016.11.003 Crossref, Google Scholar
	Carr, A.L. and Roe, M., 2016. Acarine attractants: chemoreception, bioassay, chemistry and control. Pesticide Biochemistry and Physiology 131: 60-79. https://doi.org/10.1016/j.pestbp.2015.12.009 Crossref, Google Scholar
	Carr, A.L., Sonenshine, D.E., Strider, J.B. and Roe, R.M., 2016. Evidence of female sex pheromones and characterization of the cuticular lipids of unfed, adult male versus female blacklegged ticks, Ixodes scapularis. Experimental and Applied Acarology 68: 519-538. https://doi.org/10.1007/s10493-015-0009-y Crossref, Google Scholar
	Cherry, L., 1969. Cholesterol in the cuticular wax of Boophilus microplus. Nature 222: 777. Crossref, Google Scholar
	Connat, J.L., Ducommun, J., Diehl, P.A. and Aeschlimann, A., 1986. Some aspects of the control of the gonotrophic cycle in the tick Ornithodoros moubata (Ixodoidea, Argasidae). In: Sauer, J.R. and Hair, J.A. (eds.) Morphology, physiology and behavioral biology of ticks. Ellis Horwood Limited, Chichester, UK, pp. 194-216. Google Scholar
	De Bruyne, M. and Guerin, P.M., 1994. Isolation of 2, 6-dichlorophenol from the cattle tick Boophilus microplus: receptor cell responses but no evidence for a behavioural response. Journal of Insect Physiology 40: 143-154. https://doi.org/10.1016/0022-1910(94)90086-8 Crossref, Google Scholar
	De Bruyne, M.D. and Guerin, P.M., 1998. Contact chemostimuli in the mating behaviour of the cattle tick, Boophilus microplus. Archives of Insect Biochemistry and Physiology 39: 65-80. https://doi.org/10.1002/(SICI)1520-6327(1998)39:2%3C65::AID-ARCH3%3E3.0.CO;2-W Google Scholar
	De la Vega, R., Camejo, A. and Fonseca, A.H., 2004. An automatic system to feed ticks through membranes. Review Salud Animale 26: 202-205. Google Scholar
	Dhivya, B., Latha, B.R., Raja, M.D., Sreekumar, C. and Leela, V., 2014. Control of brown dog tick, Rhipicephalus sanguineus using assembly pheromone encapsulated in natural polymer, chitosan. Experimental and Applied Acarology 63: 85-92. https://doi.org/10.1007/s10493-013-9764-9 Crossref, Google Scholar
	Diehl, P.A., Guerin, P., Vlimant, M. and Steullet, P., 1991. Biosynthesis, production site, and emission rates of aggregation-attachment pheromone in males of two Amblyomma ticks. Journal of Chemical Ecology 17: 833-847. https://doi.org/10.1007/BF01395594 Crossref, Google Scholar
	Dobrotvorsky, A.K. and Tkachev A.V., 1995. Evidence of a volatile sex pheromone in the unfed adult taiga tick Ixodes persulcatus Schulze. In: International conference on tick-borne pathogens, Kruger National Park (South Africa), 28 Aug-1 Sep 1995, pp. 372-375. Google Scholar
	Donohue, K.V, Khalil, S.M., Ross, E., Mitchell, R.D., Roe, R.M. and Sonenshine, D.E., 2009. Male engorgement factor: role in stimulating engorgement to repletion in the ixodid tick, Dermacentor variabilis. Journal of Insect Physiology 55: 909-918. https://doi.org/10.1016/j.jinsphys.2009.05.019 Crossref, Google Scholar
	Dusbábek, F., Simek, P., Jegorov, A. and Tríska, J., 1991. Identification of xanthine and hypoxanthine as components of assembly pheromone in excreta of argasid ticks. Experimental and Applied Acarology 11: 307-316. https://doi.org/10.1007/BF01202877 Crossref, Google Scholar
	Dusbábek, F., Zahradnickova, H. and Simek, P., 1998. Chemical stability of assembly pheromone of argasid ticks (Ixodoidea: Argasidae). Folia Parasitologica 45: 62-66. Google Scholar
	Dusbábek, F., Simek, P., Bouman, E.A.P., ZahradníCcková, H. and Zemek, R., 2001. The composition and effect of several semiochemicals in Ixodes ricinus (L.) (Acari: Ixodidae). In: Buczek, A. and Blaszak, C. (eds.) Proceedings of 2nd International Symposium 'Stawonogi pasozyty i nosiciele. Wyd. KGM, Lublin, Poland, pp. 33-43. Google Scholar
	El Shoura, S.M., 1987. The life cycle of Ornithodoros (Pavlovskyella) erraticus (Acari: Ixodoidea: Argasidae) in the laboratory. Journal of Medical Entomology 24: 229-34. https://doi.org/10.1093/jmedent/24.2.229 Crossref, Google Scholar
	Estrada-Peña, A., Castellá, J. and Siuda, K., 1994. Cuticular hydrocarbon composition and phenotypic variability in sympatric populations of Ixodes ricinus ticks from Poland. Experimental and Applied Acarology 18: 247-263. Crossref, Google Scholar
	Faraone, N., 2022. Host detection by ticks. Chapter 24. In: Ignell, R., Lazzari, C.R., Lorenzo, M.G. and Hill, S.R. (eds.) Sensory ecology of disease vectors. Wageningen Academic Publishers, Wageningen, the Netherlands, pp. 639653. https://doi.org/10.3920/978-90-8686-932-9_24 Google Scholar
	Ferreira, L.L., de Oliveira Filho, J.G., de Oliveira Silva, F., Ferraz, A.L. and Mascarin, G.M., 2020. Attract or repel Amblyomma sculptum ticks: screening of semiochemicals. Veterinary Parasitology 278: 109036. https://doi.org/10.1016/j.vetpar.2020.109036 Crossref, Google Scholar
	Gachoka, K.K., Ferreira, L.L., Louly, C.C.B. and Borges, L.M.F., 2011. Induction of complete courtship ritual in Amblyomma cajennense using 2, 6-dichlorophenol at female-equivalent quantities. Revista Brasileira de Parasitologia Veterinária 20: 347-350. https://doi.org/10.1590/S1984-29612011000400017 Crossref, Google Scholar
	Gachoka, K.K., Louly, C.C.B., Ferreira, L.L., Ferri, P., da Silva, A.C. and Borges, L.M.F., 2012. Chemical composition and biological characterization of extracts of male produced pheromone from Amblyomma cajennense (Fabricius, 1787) (Acari: Ixodidae) by GC/MS-SIM and behavioral responses. International Journal of Acarology 38: 445453. https://doi.org/10.1080/01647954.2012.678384 Crossref, Google Scholar
	George, J.E., 1981. The influence of aggregation pheromones on the behavior of Argas cooleyi and Ornithodoros concanensis (Acari: Ixodoidea: Argasidae). Journal of Medical Entomology 18: 129-133. https://doi.org/10.1093/jmedent/18.2.129 Crossref, Google Scholar
	Gladney, W.J., Grabbe, R.R., Ernst, S.E. and Oehler, D.D., 1974. The Gulf Coast tick: evidence of a pheromone produced by males. Journal of Medical Entomology 15: 303-6. https://doi.org/10.1093/jmedent/113.303 Google Scholar
	Gothe, R. and Kraiss, A., 1982. Zur Lokalisation der Pheromonemission und-perzeption bei Argas (Persicargas) walkerae Kaiser und Hoogstraal, 1969. Zentralblatt Veterinarmedicine Reihe B 29: 573-82. Crossref, Google Scholar
	Gothe, R. and Neitz, A.W.H., 1985. Investigation into the participation of male pheromones of Rhipicephalus evertsi evertsi during infestation. Onderstepoort Journal of Veterinary Research 52: 67-70. Google Scholar
	Gowrishankar, S., Latha, B.R., Sreekumar, C. and Leela, V., 2019. Innovative way to dispense pheromones for off-host control of Rhipicephalus sanguineus sensu lato ticks. Veterinary Parasitology 275: 108936. https://doi.org/10.1016/j.vetpar.2019.108936 Crossref, Google Scholar
	Gowrishankar, S., Latha, B.R., Sreekumar, C. and Leela, V., 2021. Solar tick trap with a pheromone lure – a stand-in approach for off-host control of Rhipicephalus sanguineus sensu lato ticks. Ticks and Tick-borne Diseases 12: 101656. https://doi.org/10.1016/j.ttbdis.2021.101656 Crossref, Google Scholar
	Graf, J.F., 1975. Ecologie et ethologie d'Ixodes ricinus L. en Suisse (Ixodoidea: Ixodidae). Cinquième note: mise en evidence d'une pheromone sexuelle chez Ixodes ricinus. Acarologia 17: 436-441. https://doi.org/10.1023/A:1016145805759 Crossref, Google Scholar
	Grenacher, S., Krober, T., Guerin, P.M. and Vlimant, M., 2001. Behavioral and chemoreceptor cell responses of the tick, Ixodes ricinus, to its own faeces and faecal constituents. Experimental and Applied Acarology 25: 641-660. https://doi.org/10.1023/A:1016145805759 Crossref, Google Scholar
	Haggart, D.A. and Davis, E.E., 1981. Neurons sensitive to 2, 6-dichlorophenol on the tarsi of the tick Amblyomma americanum (Acari: Ixodidae). Journal of Medical Entomology 18: 187-193.https://doi.org/10.2307/3280781 Crossref, Google Scholar
	Hamilton, J.G.C., 1992. The role of pheromones in tick biology. Parasitology Today 8: 130-133. https://doi.org/10.1016/0169-4758(92)90284-9 Crossref, Google Scholar
	Hamilton, J.G. and Sonenshine, D.E., 1988. Evidence for occurrence of mounting sex pheromone on body surface of female Dermacentor variabilis (Say) and Dermacentor andersoni (Stiles) (Acari: Ixodidae). Journal of Chemical Ecology 14: 401-410. https://doi.org/10.1007/BF01022555 Crossref, Google Scholar
	Hamilton, J.G.C., Papadopoulos, E., Harrison, S.J., Lloyd, C.M. and Walker, R., 1994. Evidence for a mounting sex pheromone in the brown ear tick Rhipicephalus appendiculatus, Neuman 1901 (Acari: Ixodidae). Experimental and Applied Acarology 18: 331-338. https://doi.org/10.1007/BF00116314 Crossref, Google Scholar
	Hamilton, J.G.C., Sonenshine, D.E. and Lusby, W.R., 1989. Cholesteryl oleate: mounting sex pheromone of the hard tick Dermacentor variabilis (Say) (Acari: Ixodidae). Journal of Insect Physiology 35: 873-879. https://doi.org/10.1186/1756-3305-7-182 Crossref, Google Scholar
	Hess, E. and Vlimant, M., 1986. Leg sense organs of ticks. In: Sauer, J.R. and Hair, J.A. (eds.) Morphology, physiology, and behavioral biology of ticks. John Wiley & Sons, New York, NY, USA, pp. 361-390. Google Scholar
	Kellum, D. and Berger, R.S., 1977. Relationship of the occurrence and function of 2, 6-dichlorophenol in two species of Amblyomma (Acari: Ixodidae). Journal of Medical Entomology 20: 701-705. https://doi.org/10.1093/jmedent/13.6.701 Crossref, Google Scholar
	Kelly, P.J., Lucas, H.M., Randolph, C.M., Ackerson, K., Blackburn, J.K. and Dark, M.J., 2014. Efficacy of slow-release tags impregnated with aggregation-attachment pheromone and deltamethrin for control of Amblyomma variegatum on St. Kitts, West Indies. Parasites & Vectors 7: 1-8. https://doi.org/10.1186/1756-3305-7-182 Crossref, Google Scholar
	Khalil, G.M., Sonenshine, D.E., Homsher, P.J., Dees, W.H., Carson, K.A. and Wang, V., 1983. Development, ultrastructure, and activity of the foveal glands and foveae dorsales of the camel tick, Hyalomma dromedarii (Acari: Ixodidae): 1. Primordial growth and gland formation. Journal of Medical Entomology 20: 414-423. https://doi.org/10.2307/3280781 Crossref, Google Scholar
	Khalil, G.M. 1984. Fecundity-reducing pheromone in Argas (Persicargas) arboreus (Ixodoidea: Argasidae). Parasitologia 88: 395-402. https://doi.org/10.1017/S0031182000054664 Crossref, Google Scholar
	Kiszewski, A.E., Matuschka, F.R. and Spielman, A., 2001. Mating strategies and spermiogenesis in ixodid ticks. Annual Review of Entomology 46: 167-182. https://doi.org/10.1002/ps.1293 Crossref, Google Scholar
	Kuhnert, F., 1996. Feeding of hard ticks in vitro: new perspectives for rearing and for the identification of systemic acaricides. Altex 13: 76-87. Google Scholar
	Kuhnert, F., Diehl, PA. and Guerin, P.M., 1995. The life-cycle of the bont tick Amblyomma hebraeum in vitro. International Journal for Parasitology 25: 887-896. https://doi.org/10.1093/jmedent/20.1.104 Crossref, Google Scholar
	Leahy, M.G., 1979. Pheromones of argasid ticks. Recent Advances in Acarology 2: 297-308. Crossref, Google Scholar
	Leahy, M.G. and Booth, K.S., 1983. Attraction of metastriate ticks (Acari: Ixodidae) to the sex pheromone 2, 6-dichlorophenol and to substituted phenols. Journal of Medical Entomology 20: 104-105. https://doi.org/10.1093/jmedent/20.1.104 Crossref, Google Scholar
	Leahy, M.G., Hajkova, A. and Bourchalova, J., 1981. Two female pheromones in the metastriate ticks, Hyalomma dromedarii (Acarina: Ixodidae). Acta Entomología Bohemoslovenia 78: 224-230. Google Scholar
	Leahy, M.G., Karuhize, G., Mango, C. and Galun, R., 1975a. An assembly pheromone and its perception in the tick Ornithodoros moubata (Murray) (Acari: Argasidae). Journal of Medical Entomology 12: 284-287. https://doi.org/10.1093/jmedent/12.3.284 Crossref, Google Scholar
	Leahy, M., Kovacic, A., Mannion, C. and Schulze, L., 1983. Pheromone-induced aggregation of ixodid ticks before host contact. Experientia 39: 859-860. https://doi.org/10.1007/BF01990404 Crossref, Google Scholar
	Leahy, S.M., Sternberg, S., Mango, C. and Galun, R., 1975b. Lack of specificity in assembly pheromones of soft ticks (Acari: Argasidae). Journal of Medical Entomology 12: 413-414. https://doi.org/10.1038/246515a0 Crossref, Google Scholar
	Leahy, M.G., Vandehay, R. and Galun, R., 1973. Assembly pheromones in the soft tick, Argaspersicus (Olsen). Nature 246: 515-517. https://doi.org/10.1038/246515a0 Crossref, Google Scholar
	Leonovich, S.A., 2004. Phenol and lactone receptors in the distal sensilla of the Haller’s organ in Ixodes ricinus ticks and their possible role in host perception. Experimental and Applied Acarology 32: 89-102. https://doi.org/10.1023/B:APPA.0000018200.24760.78 Crossref, Google Scholar
	Leonovich, S.A., 2020. Comparative analysis of sexual behavior in hard ticks (Ixodidae). Entomological Review 10: 122-131. https://doi.org/10.1134/S0013873820010121 Crossref, Google Scholar
	Louly, C.C.B., Silveira, D.N., Soares, S.F., Ferri, P.H., Melo, C.C. and Borges, L.M.F., 2008. More about the role of 2,6-dichlorophenol in tick courtship: identification and olfactometer bioassay in Amblyomma cajennense and Rhipicephalus sanguineus. Memórias de Instituto Oswaldo Cruz 103: 60-65. https://doi.org/10.1590/S0074-02762008000100009 Crossref, Google Scholar
	Lusby, W.R., Sonenshine, D.E., Yunker, C.E., Norval, R.A. and Burridge, M.J., 1991. Comparison of known and suspected pheromonal constituents in males of the African ticks, Amblyomma hebraeum Koch and Amblyomma variegatum (Fabricius). Experimental and Applied Acarology 13: 143-152. https://doi.org/10.1007/BF01193665 Crossref, Google Scholar
	Maranga, R.O., Hassanali, A., Kaaya, G.P. and Mueke, J.M., 2003. Attraction of Amblyomma variegatum (ticks) to the attraction-aggregation-attachment-pheromone with or without carbon dioxide. Experimental and Applied Acarology 29: 121-130. https://doi.org/10.1023/A:1024265529030 Crossref, Google Scholar
	Maranga, R.O., Hassanali, A, Kaaya, G.P. and Mueke, J.M., 2006. Performance of a prototype baited-trap in attracting and infecting the tick Amblyomma variegatum (Acari: Ixodidae) in field experiments. Experimental and Applied Acarology 38: 211-218. https://doi.org/10.1007/s10493-006-0002-6 Crossref, Google Scholar
	Maranga, R.O., Kaaya, G.P., Mueke, J.M. and Hassanali, A., 2005. Effects of combining the fungi Beauveria bassiana and Metarhizium anisopliae on the mortality of the tick Amblyomma variegatum (Ixodidae) in relation to seasonal changes. Mycopathologia 159: 527-532. https://doi.org/10.1007/s11046-005-3374-y Crossref, Google Scholar
	McDowell, P.G. and Waladde, S.M.,1986. 2, 6-dichlorophenol in the tick Rhipicephalus appendiculatus Neumann. Journal of Chemical Ecology 12: 69-81. https://doi.org/10.1007/BF01045591 Crossref, Google Scholar
	McMahon, C., Guerin, P.M. and Syed, Z., 2001. 1-Octen-3-ol isolated from bont ticks attracts Amblyomma variegatum. Journal of Chemical Ecology 27: 471-486. https://doi.org/10.1023/A:1010328720035 Crossref, Google Scholar
	Mohamed, F.S., Khalil, G.M., Marzouk, A.S. and Roshdy, M.A., 1990. Sex pheromone regulation of mating behavior in the tick Ornithodoros (Ornithodoros) savignyi (Audouin) (Acari: Argasidae). Journal of Medical Entomology 27: 288-294. https://doi.org/10.1093/jmedent/27.3.288 Crossref, Google Scholar
	Mulenga, A., 2014. Molecular biology and physiology of chemical communication. In: Sonenshine, D.E. and Roe, R.M. (eds.) Biology of ticks. Vol. 1. Oxford University Press, New York, NY, pp. 368-397. Google Scholar
	Musyoki, J.M., Osir, E.O., Kiara, H.K. and Kokwaro, E.D., 2004. Comparative studies on the infectivity of Theileria parva in ticks fed in vitro and those fed on cattle. Experimental and Applied Acarology 32: 51-67. https://doi.org/10.1023/B:APPA.0000018159.47700.e4 Crossref, Google Scholar
	Nchu, F., Maniania, N.K., Touré, A., Hassanali, A. and Eloff, J.N., 2009. The use of a semiochemical bait to enhance exposure of Amblyomma variegatum (Acari: Ixodidae) to Metarhizium anisopliae (Ascomycota: Hypocreales). Veterinary Parasitology 160: 279-284. https://doi.org/10.1016/j.vetpar.2008.11.005 Crossref, Google Scholar
	Neitz, A.W. and Gothe, R., 1984. Investigations into the volatility of female pheromones and the aggregation-inducing property of guanine in Argas (Persicargas) walkerae. Onderstepoort Journal of Veterinary Research, 51, pp.197-201. Google Scholar
	Norval, R.A. and Rechav, Y. 1979. An assembly pheromone and its perception in the tick Amblyomma variegatum (Acarina: Ixodidae). Journal of Medical Entomology 18: 507-511. https://doi.org/10.1093/jmedent/16.6.507 Crossref, Google Scholar
	Norval, R., Andrew, H.R. and Yunker, C.E., 1989a. Pheromone-mediation of host-selection in bont ticks (Amblyomma hebraeum Koch). Science 243: 364-365. https://doi.org/10.1126/science.2911745 Crossref, Google Scholar
	Norval, R.A.I., Butler, J.F. and Yunker, C.E., 1989b. Use of carbon dioxide and natural or synthetic aggregationattachment pheromone of the bont tick, Amblyomma hebraeum, to attract and trap unfed adults in the field. Experimental and Applied Acarology 7: 171-180. https://doi.org/10.1007/BF01194058 Crossref, Google Scholar
	Norval, R.A., Colborne, J., Tannock, J. and Mackenzie, P.K., 1980. The life cycle of Amblyomma tholloni Neumann, 1899 (Acarina: Ixodidae) under laboratory conditions. Veterinary Parasitology 7: 255-263. https://doi.org/10.1016/0304-4017(80)90029-1 Crossref, Google Scholar
	Norval, R.A.I., Peter, T. and Meltzer, M.I., 1992b. A comparison of the attraction of nymphs and adults of the ticks Amblyomma hebraeum and A. variegatum to carbon dioxide and the male-produced aggregation-attachment pheromone. Experimental and Applied Acarology 13: 179-186. https://doi.org/10.1007/BF01194935 Crossref, Google Scholar
	Norval, R.A.I., Peter, T., Sonenshine, D.E. and Burridge, M.J., 1992a. Responses of the ticks Amblyomma hebraeum and A. variegatum to known or potential components of the aggregation-attachment pheromone. III. Aggregation. Experimental and Applied Acarology 16: 237-245. https://doi.org/10.1007/BF01193806 Crossref, Google Scholar
	Norval, R.A.I., Peter, T., Yunker, C.E., Sonenshine, D.E. and Burridge, M.J., 1991a. Responses of the ticks Amblyomma hebraeum and A. variegatum to known or potential components of the aggregation-attachment pheromone. I. Long-range attraction. Experimental and Applied Acarology 13: 11-18. https://doi.org/10.1007/BF01268935 Crossref, Google Scholar
	Norval, R.A.I., Peter, T., Yunker, C.E., Sonenshine, D.E. and Burridge, M.J., 1991b. Responses of the ticks Amblyomma hebraeum and A. variegatum to known or potential components of the aggregation-attachment pheromone. II. Attachment stimulation. Experimental and Applied Acarology 13: 19-26. https://doi.org/10.1007/BF01268936 Crossref, Google Scholar
	Norval, R.A., Sonenshine, D.E., Allan, S.A. and Burridge, M.J., 1996. Efficacy of pheromone-acaricide-impregnated tail-tag decoys for controlling the bont tick, Amblyomma hebraeum (Acari: Ixodidae), on cattle in Zimbabwe. Experimental and Applied Acarology 20: 31-46. https://doi.org/10.1007/BF00051475 Google Scholar
	Obenchain, F.D. 1984. Behavioural interactions between the sexes and aspects of species specificity pheromone mediated aggregation and attachment in Amblyomma. In: Griffith, D.A. and Bowman, C.E. (eds.) Acarology, vol 1. Ellis Horwood, Chichester, UK, pp. 387-392. Google Scholar
	Oliver, J.D., Lynn, G.E., Burkhardt, N.Y., Price, L.D., Nelson, C.M., Kurtti, T.J. and Munderloh, U.G., 2016. Infection of immature Ixodes scapularis (Acari: Ixodidae) by membrane feeding. Journal of Medical Entomology 53: 409415. https://doi.org/10.1093/jme/tjv241 Crossref, Google Scholar
	Otieno, D.A., Hassanali, A., Obenchain, F.D., Sternberg, A. and Galun, R., 1985. Identification of guanine as an assembly pheromone of ticks. International Journal of Tropical Insect Science 6: 667-70 https://doi.org/10.1017/S1742758400002873 Crossref, Google Scholar
	Pappas, P.J. and Oliver Jr, J.H., 1972. Reproduction in ticks (Acari: Ixodoidea) 2. Analysis of the stimulus for rapid and complete feeding of female Dermacentor variabilis (Say). Journal of Medical Entomology 9: 47-50. https://doi.org/10.1093/jmedent/9.1.47 Crossref, Google Scholar
	Petney, T.N. and Andrews, R.H., 1982. The influence of similar aggregation pheromones on the microhabitat choice of two parapatric species of reptile tick (Acari: Ixodidae). Oecologia 55: 364-368. https://doi.org/10.1007/BF00376924 Crossref, Google Scholar
	Petney, T.N. and Bull, C.M., 1981. A non-specific aggregation pheromone in two Australian reptile ticks. Animal Behavior 29: 181-185. https://doi.org/10.1016/S0003-3472(81)80164-9 Crossref, Google Scholar
	Phillips, J. and Sonenshine, D.E., 1993. Role of the male claw sensilla in perception of female mounting sex pheromone in Dermacentor variabilis, Dermacentor andersoni and Amblyomma americanum. Experimental and Applied Acarology 17: 631-653. https://doi.org/10.1007/BF00058504 Crossref, Google Scholar
	Pitts, R.J., Ibarra Bouzada, L.M.E. and Guerenstein, PG., 2022. Comparative morphology of the peripheral olfactory system of disease vector arthropods. Chapter 2. In: Ignell, R., Lazzari, C.R., Lorenzo, M.G. and Hill, S.R. (eds.) Sensory ecology of disease vectors. Wageningen Academic Publishers, Wageningen, the Netherlands, pp. 29-70. https://doi.org/10.3920/978-90-8686-932-9_2 Google Scholar
	Price, T.L., Sonenshine, D.E., Norval, R.A.I., Yunker, C.E. and Burridge, M.J., 1994. Pheromonal composition of two species of African Amblyomma ticks: similarities, differences and possible species specific components. Experimental and Applied Acarology 18: 37-50. https://doi.org/10.1007/BF00051401 Crossref, Google Scholar
	Ranju, R.S., Bhaskaran, R.L. and Leela, V., 2018. Exploiting assembly pheromone for the control of ixodid ticks. Acarologia 58: 471-482.https://dx.doi.org/10.24349/acarologia/20184254 Crossref, Google Scholar
	Ranju, R.S., Latha, B.R., Leela, V. and Basith, S.A. 2012. Effect of attractant sex pheromone on immature larval stages of ixodid tick species. Journal of Parasitic Diseases 36: 155-158. https://doi.org/10.1007/s12639-012-0140-1 Crossref, Google Scholar
	Ranju, R.S., Latha, B.R., Leela, V. and Basith, S.A. 2013. Field trials to attract questing stages of brown dog tick, Rhipicephalus sanguineus using tick pheromone-acaricide complex. Journal of Parasitic Diseases 37: 84-87. https://doi.org/10.1007/s12639-012-0136-x Crossref, Google Scholar
	Rechav, Y. and Whitehead, G.N., 1978. Field trials with pheromone-acaricide mixtures for control of Amblyomma hebraeum. Journal of Economic Entomology 71: 149-151. https://doi.org/10.1093/jee/71.1.149 Crossref, Google Scholar
	Rechav, Y., Goldberg, M. and Fielden, L.J., 1997. Evidence for attachment pheromones in the Cayenne tick (Acari: Ixodidae). Journal of Medical Entomology 34: 234-237. https://doi.org/10.1093/jmedent/34.2.234 Crossref, Google Scholar
	Rechav, Y., Parolis, H., Whitehead, G.B. and Knight, M.M., 1977. Evidence for an assembly pheromone (s) produced by males of the bont tick, Amblyomma hebraeum (Acarina: Ixodidae). Journal of Medical Entomology 14: 71-78. https://doi.org/10.1093/jmedent/14.1.71 Crossref, Google Scholar
	Rechav, Y., Drey, C., Fielden, L.J. and Goldberg, M., 2000. Production of pheromones by artificially fed males of the tick Amblyomma maculatum (Acari: Ixodidae). Journal of Medical Entomology 37: 761-765. https://doi.org/10.1603/0022-2585-37.5.761 Crossref, Google Scholar
	Renthal, R., Lohmeyer, K., Borges, L.M. and de León, A.A., 2019. Surface lipidome of the lone star tick, Amblyomma americanum, provides leads on semiochemicals and lipid metabolism. Ticks and Tick-borne Diseases 10: 138145. https://doi.org/10.1016/j.ttbdis.2018.09.009 Crossref, Google Scholar
	Schlein, Y. and Gunders, A.E., 1981. Pheromone of Ornithodoros spp. (Argasidae) in the coxal fluid of female ticks. Parasitology 82: 467-471. https://doi.org/10.1017/S0031182000066993 Crossref, Google Scholar
	Schöni, R., Hess, E., Blum, W. and Ramstein, K., 1984. The aggregation-attachment pheromone of the tropical bont tick Amblyomma variegatum Fabricius (Acari, Ixodidae): isolation, identification and action of its components. Journal of Insect Physiology 30: 613-618. https://doi.org/10.1016/0022-1910(84)90045-3 Crossref, Google Scholar
	Silverstein, R.M., West, J.R., Sonenshine, D.E. and Khalil, G.M., 1983. Occurrence of 2, 6-dichlorophenol in hard ticks, Hyalomma dromedarii and Hyalomma anatolicum excavatum, and its role in mating. Journal of Chemical Ecology 9: 1543-1549. https://doi.org/10.1007/BF00988420 Crossref, Google Scholar
	Sleeba, S.B., Teel, P.D., Longnecker, M.T. and Strey, O.F., 2010. Host selection by questing female Amblyomma maculatum Koch, to cattle with feeding male ticks in southern Texas. Veterinary Parasitology 172: 105-108. https://doi.org/10.1016/j.vetpar.2010.04.031 Crossref, Google Scholar
	Sobbhy, H., Aggour, M.G., Sonenshine, D.E. and Burridge, M.J., 1994. Cholesteryl esters on the body surfaces of the camel tick, Hyalomma dromedarii (Koch, 1844) and the brown dog tick, Rhipicephalus sanguineus (Latreille, 1806). Experimental and Applied Acarology 18: 265-280. https://doi.org/10.1007/BF00132316 Crossref, Google Scholar
	Soares, S.F. and Borges, L.M.F., 2012. Electrophysiological responses of the olfactory receptors of the tick Amblyomma cajennense (Acari: Ixodidae) to host-related and tick pheromone-related synthetic compounds. Acta Tropica 124: 192-198. https://doi.org/10.1016/j.actatropica.2012.08.007 Crossref, Google Scholar
	Sonenshine, D.E., 1985. Pheromones and other semiochemicals of the Acari. Annual Review of Entomology 30: 1-28. Sonenshine, D.E., 1991. Biology of Ticks. Vol. 1. Oxford University Press, New York, New York, USA, pp. 447. Sonenshine, D.E., 2004. Pheromones and other semiochemicals of ticks and their use in tick control. Parasitology 129: S405-425. https://doi.org/10.1017/S003118200400486X Crossref, Google Scholar
	Sonenshine, D.E., 2006. Tick pheromones and their use in tick control. Annual Review of Entomology 51: 557-580. https://doi.org/10.1146/annurev.ento.51.110104.151150 Crossref, Google Scholar
	Sonenshine, D.E., 2008. Pheromones and other semiochemicals of ticks and their use for tick control. In: Bowman, A.S. and Nuttall, P.A. (eds.) Ticks: biology, disease and control. Cambridge University Press, Cambridge, UK, pp. 405-425. Crossref, Google Scholar
	Sonenshine, D.E. and Hamilton, J.G.C., 1989. Methods and apparatus for controlling arthropod populations. US Patent No. 4884361. Google Scholar
	Sonenshine, D.E., Adams, T., Allan, S.A., McLaughlin, J. and Webster, F.X., 2003. Chemical composition of some components of the arrestment pheromone of the black-legged tick, Ixodes scapularis (Acari: Ixodidae) and their use in tick control. Journal of Medical Entomology 40: 849-859. https://doi.org/10.1603/0022-2585-40.6.849 Crossref, Google Scholar
	Sonenshine, D.E., Allan, S.A., Peter, T.F., McDaniel, R. and Burridge, M.J., 2000. Does geographic range affect the attractant-aggregation-attachment pheromone of the tropical bont tick, Amblyomma variegatum? Experimental and Applied Acarology 24: 283-299. https://doi.org/10.1023/A:1006479118160 Crossref, Google Scholar
	Sonenshine, D.E., Bissinger, B.W., Egekwu, N., Donohue, K.V., Khalil, S.M. and Roe, R.M., 2011. First transcriptome of the testis-vas deferens-male accessory gland and proteome of the spermatophore from Dermacentor variabilis (Acari: Ixodidae). PloS ONE 6: e24711. https://doi.org/10.1371/journal.pone.0024711 Crossref, Google Scholar
	Sonenshine, D.E., Homsher, P.J., Carson, K.A. and Wang, V.D., 1984. Evidence of the role of the cheliceral digits in the perception of genital sex pheromones during mating in the American dog tick, Dermacentor variabilis (Acari: Ixodidae). Journal of Medical Entomology 21: 296-306. https://doi.org/10.1093/jmedent/21.3.296 Crossref, Google Scholar
	Sonenshine, D.E., Khalil, G.M., Homsher, P.J. and Mason, S.N., 1982. Dermacentor variabilis and Dermacentor andersoni: genital sex pheromones. Experimental Parasitology 54: 317-330. https://doi.org/10.1016/0014-4894(82)90041-8 Crossref, Google Scholar
	Sonenshine, D.E., Silverstein, R.M., Brossut, R., Davis, E.E., Taylor, D., Carson, K.A., Homsher, P.J. and Wang, V.B., 1985a. Genital sex pheromones of ixodid ticks: 1. Evidence of occurrence in anterior reproductive tract of American dog tick, Dermacentor variabilis (Say)(Acari: Ixodidae). Journal of Chemical Ecology 11: 1669-1694. https://doi.org/10.1007/BF01012120 Crossref, Google Scholar
	Sonenshine, D.E., Silverstein, R.M., Plummer, E., West, J.R. and McCullough, T., 1976. 2, 6-Dichlorophenol, the sex pheromone of the Rocky Mountain wood tick, Dermacentor andersoni Stiles and the American dog tick, Dermacentor variabilis (Say). Journal of Chemical Ecology 2: 201-209. https://doi.org/10.1007/BF00987743 Crossref, Google Scholar
	Sonenshine, D.E., Taylor, D. and Corrigan, G., 1985b. Studies to evaluate the effectiveness of sex pheromone-impregnated formulations for control of populations of the American dog tick, Dermacentor variabilis (Say) (Acari: Ixodidae). Experimental and Applied Acarology 1: 23-34. Crossref, Google Scholar
	Steullet, P and Guerin, PM. 1994. Identification of vertebrate volatiles stimulating olfactory receptors on tarsus I of the tick Amblyomma variegatum Fabricius (Ixodidae). II. Receptors outside the Haller’s organ capsule. Journal of Comparative Physiology A 174: 3947. https://doi.org/10.1007/BF00192003 Crossref, Google Scholar
	Tajeri, S. and Razmi, G.R., 2011. Hyalomma anatolicum anatolicum and Hyalomma dromedarii (Acari: Ixodidae) imbibe bovine blood in vitro by utilizing an artificial feeding system. Veterinary Parasitology 180: 332-335. https://doi.org/10.1016/j.vetpar.2011.03.014 Crossref, Google Scholar
	Taylor, D., Phillips, J.S., Allan, S.A. and Sonenshine, D.E., 1987. Absence of assembly pheromones in the hard ticks Dermacentor variabilis and Dermacentor andersoni (Acari: Ixodidae). Journal of Medical Entomology 24: 628-32. https://doi.org/10.1093/jmedent/24.6.628 Crossref, Google Scholar
	Taylor, D., Phillips, J.S., Sonenshine, D.E. and Hanson, F.E., 1991. Ecdysteroids as a component of the genital sex pheromone in two species of hard ticks, Dermacentor variabilis and Dermacentor andersoni (Acari: Ixodidae). Experimental and Applied Acarology 12: 275-296. https://doi.org/10.1007/BF01193473 Crossref, Google Scholar
	Tkachev, A.V., Dobrotvorsky, A.K., Vjalkov, A.I. and Morozov, S.V., 2000. Chemical composition of lipophylic compounds from the body surface of unfed adult Ixodes persulcatus ticks (Acari: Ixodidae). Experimental and Applied Acarology 24: 145-158. https://doi.org/10.1023/A:1006430323587 Crossref, Google Scholar
	Treverrow, N.L., Stone, B.F. and Cowie, M., 1977 Aggregation pheromones in two Australian hard ticks, Ixodes holocyclus and Aponomma concolor. Experientia 33: 680-682. https://doi.org/10.1007/BF01946573 Crossref, Google Scholar
	Uspensky, I.V. and Emel’yanova, O.Y., 1980. Existence of pheromone associations in ticks of the genus Ixodes. Zoología Zh. 59: 699-704 (Engl. transl. T1447, NAMRU, Cairo) Google Scholar
	Waladde, S.M., Young, A.S. and Morzaria, S.P., 1996. Artificial feeding of ixodid ticks. Parasitology Today 12: 272278. https://doi.org/10.1016/0169-4758(96)10027-2 Crossref, Google Scholar
	Waladde, S.M., Young, A.S., Mwaura, S.N. and Mwakima, F.N., 1993. Transmission of Theileria parva to cattle by Rhipicephalus appendiculatus adults fed as nymphae in vitro on infected blood through an artificial membrane. Parasitology 107: 249-256. https://doi.org/10.1111/j.1749-6632.1996.tb53527.x Crossref, Google Scholar
	Walker, A.R., Lloyd, C.M., Mcguire, K., Harrison, S.J. and Hamilton, J.G.C., 2014. Integumental glands of the tick Rhipicephalus appendiculatus (Acari: Ixodidae) as potential producers of semiochemicals. Journal of Medical Entomology 33: 743-759. https://doi.org/10.1093/jmedent/33.5.743 Crossref, Google Scholar
	Wang, H., Paesen, G.C., Nuttall, P.A. and Barbour, A.G., 1998. Male ticks help their mates to feed. Nature 391: 753754. https://doi.org/10.1038/35773 Crossref, Google Scholar
	Weiss, B.L. and Kaufman, W.R., 2004. Two feeding-induced proteins from the male gonad trigger engorgement of the female tick Amblyomma hebraeum. Proceedings of the National Academy of Sciences of the USA 101: 5874-5879. https://doi.org//10.1073/pnas.0307529101 Crossref, Google Scholar
	Yoder, J.A. and Stevens, B.W., 2000. Attraction of immature stages of the American dog tick (Dermacentor variabilis) to 2, 6-dichlorophenol. Experimental and Applied Acarology 24: 159-164. https://doi.org/10.1023/A:1006419203251 Crossref, Google Scholar
	Yoder, J.A., Domingus, J.L. and Luerman, G.C., 2003. Uric acid-induced arrestment as a possible bird host cue (kairomone) in nymphs and adults of the lone star tick, Amblyomma americanum (L.). International Journal of Acarology 29: 399-402. https://doi.org/10.1080/01647950308684357 Crossref, Google Scholar
	Yoder, J.A., Hanson, P.E., Pizzuli, J.L., Sanders, C.I. and Domingus, J.L., 2009. Pheromone production and its relationship to water conservation: studies on a trichlorophenol in the American dog tick, Dermacentor variabilis (Acari: Ixodidae). International Journal of Acarology 28: 379-385. https://doi.org/10.1080/01647950208684314 Crossref, Google Scholar
	Yoder, J.A., Hanson, PE., Sanders, C.I. and Burke, W.J., 2002. On the role of 2,6-dichlorophenol as a tick sex pheromone (Acari: Ixodidae), International Journal of Acarology 28: 49-54. https://doi.org/10.1080/01647950208684280 Crossref, Google Scholar
	Yoder, J.A., Rausch, B.A., Jajack, A.J., Tomko, P.M., Gribbins, K.M. and Benoit, J.B., 2013. Snakes produce kairomones that induce aggregation of unfed larval blacklegged ticks Ixodes scapularis (Acari: Ixodidae). International Journal of Acarology 39: 502-506. https://doi.org/10.1080/01647954.2013.837507 Crossref, Google Scholar
	Young, A.S., Waladde, S.M. and Morzaria, S.P., 1996. Artificial feeding systems for ixodid ticks as a tool for study of pathogen transmission. Annals of the New York Academy of Science 791: 211-218. https://doi.org/10.1111/j.1749-6632.1996.tb53527.x Crossref, Google Scholar
	Zemek, R., Bouman, E., Socha, R. and Dusbábek, F., 2002. The effect of feeding status on sexual attractiveness of Ixodes ricinus (Acari: Ixodidae) females. Experimental and Applied Acarology 27: 137-149. https://doi.org/10.1023/A:1021511505415 Crossref, Google Scholar
	Zemek, R., Bouman, E.A.P and Dusbábek, F., 2007. The influence of conspecific chemical cues on walking behavior of Ixodes ricinus males. Experimental and Applied Acarology 41: 255-265. https://doi.org/10.1007/s10493-007-9067-0 Crossref, Google Scholar
	Ziv, M., Sonenshine, D.E., Silverstein, R.M., West, J.R. and Gingher, K.H., 1981. Use of sex pheromone, 2, 6-dichlorophenol, to disrupt mating by American dog tick Dermacentor variabilis (Say). Journal of Chemical Ecology 7: 829-840. https://doi.org/10.1007/BF00992381 Crossref, Google Scholar